First Report of Dinosaur Footprints from Madagascar: Two Tracksites from the Middle Jurassic Bemaraha Formation

Here we report the first record of tetrapod tracks from Madagascar. We document two localities yielding dinosaur footprints, both within the Middle Jurassic Bemaraha Formation (Morondava Basin) in western Madagascar. The Sahalaly River tracksite yielded a single trackway belonging to a quadrupedal dinosaur; probably a sauropod, but a closer determination is hindered by bad preservation. In contrast, the Tsiandro tracksite yielded numerous well-preserved theropod tracks that allow some inference about the behavior of the trackmakers.